Jan 8, 2018
INTRODUCTION
Images below are From:
http://bcmushrooms.forrex.org/ntfp/pages/introduction.html
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In
1999, Wills and Lipsey identified a group of provincial non-timber
forest products (NTFP) and services, including wild mushrooms, that can
be commercially harvested from forests and other habitats for culinary
or reputed nutriceutical purposes. The wild mushroom harvest alone contributes millions of dollars to the provincial economy. The pine mushroom (Tricholoma magnivelare),
is by far the most valuable commercially harvested mushroom. Other
valuable edible mushrooms are the chanterelles, morels, lobsters,
boletes, cauliflowers and hedgehogs. Pine mushrooms are exported
exclusively to Japan, while chanterelles and other wild mushrooms are
exported primarily to Europe and other parts of North America. Medical
research, predominantly from Asia, suggests that many compounds derived
from mushrooms prevent illnesses or boost human health. The demand for
wild mushrooms is, therefore, expected to increase.
Biological,
taxonomical and ecological information on mushrooms that are currently
(or could potentially be) harvested commercially is scattered throughout
various scientific and technical publications. Developing an
understanding of these wild mushrooms can be an arduous task for someone
with limited mycological training. People with minimal mycological
experience who are involved in the wild mushroom harvest -- pickers,
sellers, buyers, foresters, managers, ecologists and naturalists --
require a concise, user-friendly mushroom information source. This web
site seeks to bring together such scientific and technical information
on our province's edible wild mushrooms.
Mushroom Biology
A
mushroom is the above ground fruiting body of a fungus. Unlike plants,
fungi do not manufacture their own food but must absorb their food from
an external source. They obtain their food in three ways, as:
- saprophytes that live off dead organic material;
- symbionts that form mutually beneficial; relationships with living organisms
- parasites or pathogens that infect live off living organisms.
Many forest mushrooms
form symbiotic relationships known as mycorrhizae with tree roots.
Mycorrhizal mushrooms are critical to the health of their hosts.
Saprophytes included on this web site decompose organic matter such as
decaying wood, forest litter and other plant material. Parasitic
mushrooms included here are pathogens that either cause butt or root
rots of conifer and deciduous trees or infect and actually 'feed off'
other mushrooms.
All
mushrooms have at least two parts to their life cycle: (1) a vegetative
stage dedicated to growth and (2) a reproductive stage (fruiting
bodies) dedicated to spore production. The vegetative stage is composed
of filamentous threads known as hyphae through which water and nutrients
move throughout the fungus. Under ideal conditions, hyphae grow rapidly
and form a complex intricate network known as the mycelium. When
environmental conditions are suitable, the mycelium forms fruiting
bodies that come in a variety of shapes, sizes, colours, tastes and
odours. Mushrooms are fruiting bodies formed above ground while truffles
form fruiting bodies below ground. Typical mushrooms have caps (pilei),
gills (lamellae) and stalks (stipes) that vary greatly in form.
Mushrooms belonging to the Basidiomycetes produce spores on sexual
structures called basidia that are found underneath the cap on gills,
folds or veins, pores or tubes, or in spines or teeth (see illustration
below). The fertile layer or tissue is called the hymenium. Mushrooms
belonging to the Ascomycetes, such as morels, produce their spores
inside sac-like structures called asci. Spores are dispersed to new
habitats by wind, water, air currents and animals that eat the fruiting
bodies. Basidia, asci and spores are microscopic structures. Spore
shape, size and chemical reactivity are important characters used to
identify mushrooms.
Mushroom Names
We
include both scientific and common names of mushrooms. Scientific (or
Latin) names are used because they are the same no matter what the
spoken language may be and their binomial name can help indicate
relationships among similar species. Some morphological species are
difficult to differentiate und field conditions and molecular analysis
is increasingly being used to separate species. These tests are
revealing unexpected relationships among traditional species groups. As a
result, species concepts are changing, with species transferred between
genera, and genera between families. New genera and families are being
erected, even as previously used family and generic names are
disappearing. With each transfer, the scientific name changes. Synonyms
are previously used names that reflect where earlier scientists have
classified the mushroom. For instance, the pine mushroom of B.C. was at
one time called Armillaria ponderosa and Tricholoma ponderosa, but taxonomists have determined that pine mushrooms belong to the genus Tricholoma and nomenclatural rules dictate that the first named species –– in this case magnivelare –– is the valid name and must be used. Each mushroom’s description includes its synonyms.
We
also include the etymology or history of the scientific name for each
species because it can help us to remember the Latin name. For instance,
the Latin root ‘lact’ means ‘milk’ in English, which helps us remember that any mushroom name including “lact” in its name will probably exude a milk–like liquid when cut.
Common
names reflect regional and local preferences. Over time, various people
have coined vernacular names for a variety of different reasons. Some
common names stick, others don’t, and rarely do such names reflect
similarities or affinities among or within groups. Nonetheless, they can
be useful so we present both the preferred common name as well as the
more common Latin synonyms.
Mushroom Identification
No
web site can substitute for training in mushroom identification. Some
mushrooms in B.C. are deadly poisonous, others are moderately poisonous
and others are safe to eat for most people. Some edible mushrooms cause
negative reactions in some people. So it is important to never eat
mushrooms that haven’t been correctly identified.
All
species descriptions are based on fresh specimens and are organized by
features of the cap, hymenium and stalk. Pertinent micro-features such
as spore color, shape, and size, shape of basidia, presence of clamp
connections, etc., are also included.
Macroscopic Features
Cap (Pileus)
Diameter
of both the button and mature caps are measured. The variations among
the most common cap shapes are presented below. Surface texture is
described as being smooth, furry, fibrillose, scaly, sticky, slimy,
cracking, or splitting (especially towards the margins). Colour and any
colour changes should be noted and described using a standard colour
chart. After mature and immature caps are cut, their texture, colour,
bruising colour changes, and width at the stalk apex as well as at the
base should be described.
|
Taste and Odour
Tastes
and odours of fresh mushrooms can be distinctive and help identify a
mushroom, so taste and odours of fresh specimens should be noted. Be aware that some mushrooms are deadly poisonous, so do not swallow tissues of unknown mushrooms. [Taste and “spit”!]
Gills, Folds, Veins, Pores, Tubes or Spines (Hymenium)
The
gills, folds, pores or spines are attached to the stalk in a variety of
ways (see illustrations below). Spacing, depth, nature of the edge and
branching pattern are all described, as are the colour of gills in both
button and mature mushrooms as well as any colour changes.
Stalk (Stipe)
Not
all mushrooms have stalks, but if they do, the length (from apex to
base) as well as the width (both at the apex and and at the base) are
measured. Other features noted include the stalk’s shape, its surface
texture, its colour, and the texture and colour of the context (the
flesh or internal tissue). If a ring (annulus) and basal cup (volva) are
present, they are described.
|
|
Microscopic Features
Spores
Spore
prints are key aids in identifying mushrooms. Separate a mature cap
from the stalk and place it on white or black paper with the hymenium
facing down, so that spores collect on the paper. Place the cap and
paper inside a lightly covered container and leave outside in ambient
temperatures for 12–24 hours. Note the spore print colour (without
magnification) to help identify and classify the mushroom. To examine
the spores microscopically, place the hymenium tissue on a microscope
slide in water or Melzer’s reagent and cover with a glass cover slip
before examining under
the microscope. Measure and examine the spores. Spores are placed in
Melzer’s reagent because the spores’ reaction helps in mushroom
identification. Sometimes, to identify unknown specimens, additional
reagents such as cotton blue are used as an aid to precise
identification.
Other Features
Clamp
connections, which are connecting cells between two adjoining hyphal
cells, occur only in fungi belonging to the Basidiomycetes. The presence
or absence of clamp connections can assist in identifying mushrooms in
this group.
The length of the basidia or asci and the number of spores produced by them also help to identify some mushrooms.
Hymenial
cystidia are sterile cells interspersed among the basidia in the
hymenium, and their presence or absence can also be important. When they
are present, their shape and size are very helpful in indentifying
mushrooms.
Mushroom Habitats and Habits
Habitat
includes the ecological setting in which a mushroom is most likely to
be found. This includes the host trees or host substrates as well as the
surrounding vegetation. Some mushrooms can colonize a wide range of
substrates such as dead wood or twigs or hosts, while others are very
specific to certain substrates or hosts. For instance, Boletus mirabilis, the admirable bolete, forms ectomycorrhizae only with western hemlock, while Tricholoma magnivelare,
the pine mushroom, colonizes a variety of conifer trees. Likewise, some
saprophytic mushrooms decompose a wide range of woody hosts, while
others decompose very specific woody hosts. Some saprophytes decompose
forest litter, while others decompose organic matter in pastures, lawns
or disturbed areas.
The
vast majority of B.C. NTFP mushrooms fruit in the autumn, although some
fruit only in the spring. Others may fruit throughout the year, weather
permitting. It is important to know in which season a mushroom fruits
when hunting for a specific mushroom. Fruiting itself is triggered by
such environmental factors as moisture and temperature. Fruiting
patterns of specific mushrooms vary, as some fruit singly, others fruit
in troops and still others form clusters. On this website, we provide
the fruiting habit of each mushroom.
Distribution Maps
The
distribution map for each species is based on collection areas
confirmed from personal and herbarium collections and published reports.
There are undoubtedly parts of the province where some species occur
but which are not yet included in these maps. As more data are added,
these distribution maps will more accurately reflect the entire range
and distribution of each species.
Interesting Facts
Mushrooms
have been part of human civilizations for centuries. One of the
earliest records of mushroom use was the case of Otzi, the Tyrolean
Iceman, who was frozen in ice but not discovered until 1991 in the
Italian Alps. Otzi lived 5,000 years ago and was carrying three
different mushrooms when he died. The fungi had laxative and antibiotic
properties, and scientists speculate that Otzi used the fungi to cure
his stomach parasites. In Asia, mushrooms have been used for centuries
for culinary, medicinal, nutriceutical and aphrodisiacal purposes.
Ancient Greeks and Romans welcomed thunderstorms because they believed
that mushrooms appeared after lightning. There is a plethora of
beneficial and unusual uses attributed to certain mushrooms, some of
which we include in our treatment of individual species. We also provide
unusual items of interest about each species. Some fungi have a
livelier or longer history than others, so for some species there is an
extensive section on interesting facts.
Disclaimer
No
single test can determine the culinary, beneficial or toxic properties
of a specific mushroom. As some mushrooms can be deadly poisonous, we
emphasize that the only safe means of separating edible mushrooms from
poisonous ones is the positive identification by an expert.
For
over twenty centuries, Chinese doctors have used mushrooms and other
fungi to cure many types of human diseases. These medicinal mushrooms
have also been credited with the ability to promote health and boost the
immune system. However, Western medicine is still struggling to confirm
Eastern medicinal uses of mushrooms.
Many
mushrooms produce both beneficial and toxic compounds; the same
compound can have beneficial or detrimental effects depending on the
amount consumed, how the mushroom is prepared, what other foods or
beverages are consumed with it, and whether an individual is
over-senstive to the active compounds. We neither confirm nor deny the
actions attributed to individual mushroom compounds. Although we are
aware of many incidences where people attribute their well being to a
specific mushroom, we try to avoid repeating anecdotal evidence and cite
only published scientific research with respect to the healing or other
beneficial properties attributed to each species. We do direct the
reader to published reports of the benefits of specific mushrooms after
each species description.
Reference
Wills,
R.M., and R.G. Lipsey, 1999. An economic strategy to develop Non-Timber
Forest Products and Services in British Columbia. Forest Renewal BC
Project No. PA97538-ORE. Final Report.
|
|
|
|
|
In
1999, Wills and Lipsey identified a group of provincial non-timber
forest products (NTFP) and services, including wild mushrooms, that can
be commercially harvested from forests and other habitats for culinary
or reputed nutriceutical purposes. The wild mushroom harvest alone contributes millions of dollars to the provincial economy. The pine mushroom (Tricholoma magnivelare),
is by far the most valuable commercially harvested mushroom. Other
valuable edible mushrooms are the chanterelles, morels, lobsters,
boletes, cauliflowers and hedgehogs. Pine mushrooms are exported
exclusively to Japan, while chanterelles and other wild mushrooms are
exported primarily to Europe and other parts of North America. Medical
research, predominantly from Asia, suggests that many compounds derived
from mushrooms prevent illnesses or boost human health. The demand for
wild mushrooms is, therefore, expected to increase.
Biological,
taxonomical and ecological information on mushrooms that are currently
(or could potentially be) harvested commercially is scattered throughout
various scientific and technical publications. Developing an
understanding of these wild mushrooms can be an arduous task for someone
with limited mycological training. People with minimal mycological
experience who are involved in the wild mushroom harvest -- pickers,
sellers, buyers, foresters, managers, ecologists and naturalists --
require a concise, user-friendly mushroom information source. This web
site seeks to bring together such scientific and technical information
on our province's edible wild mushrooms.
Mushroom Biology
A
mushroom is the above ground fruiting body of a fungus. Unlike plants,
fungi do not manufacture their own food but must absorb their food from
an external source. They obtain their food in three ways, as:
- saprophytes that live off dead organic material;
- symbionts that form mutually beneficial; relationships with living organisms
- parasites or pathogens that infect live off living organisms.
Many forest mushrooms
form symbiotic relationships known as mycorrhizae with tree roots.
Mycorrhizal mushrooms are critical to the health of their hosts.
Saprophytes included on this web site decompose organic matter such as
decaying wood, forest litter and other plant material. Parasitic
mushrooms included here are pathogens that either cause butt or root
rots of conifer and deciduous trees or infect and actually 'feed off'
other mushrooms.
All
mushrooms have at least two parts to their life cycle: (1) a vegetative
stage dedicated to growth and (2) a reproductive stage (fruiting
bodies) dedicated to spore production. The vegetative stage is composed
of filamentous threads known as hyphae through which water and nutrients
move throughout the fungus. Under ideal conditions, hyphae grow rapidly
and form a complex intricate network known as the mycelium. When
environmental conditions are suitable, the mycelium forms fruiting
bodies that come in a variety of shapes, sizes, colours, tastes and
odours. Mushrooms are fruiting bodies formed above ground while truffles
form fruiting bodies below ground. Typical mushrooms have caps (pilei),
gills (lamellae) and stalks (stipes) that vary greatly in form.
Mushrooms belonging to the Basidiomycetes produce spores on sexual
structures called basidia that are found underneath the cap on gills,
folds or veins, pores or tubes, or in spines or teeth (see illustration
below). The fertile layer or tissue is called the hymenium. Mushrooms
belonging to the Ascomycetes, such as morels, produce their spores
inside sac-like structures called asci. Spores are dispersed to new
habitats by wind, water, air currents and animals that eat the fruiting
bodies. Basidia, asci and spores are microscopic structures. Spore
shape, size and chemical reactivity are important characters used to
identify mushrooms.
Mushroom Names
We
include both scientific and common names of mushrooms. Scientific (or
Latin) names are used because they are the same no matter what the
spoken language may be and their binomial name can help indicate
relationships among similar species. Some morphological species are
difficult to differentiate und field conditions and molecular analysis
is increasingly being used to separate species. These tests are
revealing unexpected relationships among traditional species groups. As a
result, species concepts are changing, with species transferred between
genera, and genera between families. New genera and families are being
erected, even as previously used family and generic names are
disappearing. With each transfer, the scientific name changes. Synonyms
are previously used names that reflect where earlier scientists have
classified the mushroom. For instance, the pine mushroom of B.C. was at
one time called Armillaria ponderosa and Tricholoma ponderosa, but taxonomists have determined that pine mushrooms belong to the genus Tricholoma and nomenclatural rules dictate that the first named species –– in this case magnivelare –– is the valid name and must be used. Each mushroom’s description includes its synonyms.
We
also include the etymology or history of the scientific name for each
species because it can help us to remember the Latin name. For instance,
the Latin root ‘lact’ means ‘milk’ in English, which helps us remember that any mushroom name including “lact” in its name will probably exude a milk–like liquid when cut.
Common
names reflect regional and local preferences. Over time, various people
have coined vernacular names for a variety of different reasons. Some
common names stick, others don’t, and rarely do such names reflect
similarities or affinities among or within groups. Nonetheless, they can
be useful so we present both the preferred common name as well as the
more common Latin synonyms.
Mushroom Identification
No
web site can substitute for training in mushroom identification. Some
mushrooms in B.C. are deadly poisonous, others are moderately poisonous
and others are safe to eat for most people. Some edible mushrooms cause
negative reactions in some people. So it is important to never eat
mushrooms that haven’t been correctly identified.
All
species descriptions are based on fresh specimens and are organized by
features of the cap, hymenium and stalk. Pertinent micro-features such
as spore color, shape, and size, shape of basidia, presence of clamp
connections, etc., are also included.
Macroscopic Features
Cap (Pileus)
Diameter
of both the button and mature caps are measured. The variations among
the most common cap shapes are presented below. Surface texture is
described as being smooth, furry, fibrillose, scaly, sticky, slimy,
cracking, or splitting (especially towards the margins). Colour and any
colour changes should be noted and described using a standard colour
chart. After mature and immature caps are cut, their texture, colour,
bruising colour changes, and width at the stalk apex as well as at the
base should be described.
|
Taste and Odour
Tastes
and odours of fresh mushrooms can be distinctive and help identify a
mushroom, so taste and odours of fresh specimens should be noted. Be aware that some mushrooms are deadly poisonous, so do not swallow tissues of unknown mushrooms. [Taste and “spit”!]
Gills, Folds, Veins, Pores, Tubes or Spines (Hymenium)
The
gills, folds, pores or spines are attached to the stalk in a variety of
ways (see illustrations below). Spacing, depth, nature of the edge and
branching pattern are all described, as are the colour of gills in both
button and mature mushrooms as well as any colour changes.
Stalk (Stipe)
Not
all mushrooms have stalks, but if they do, the length (from apex to
base) as well as the width (both at the apex and and at the base) are
measured. Other features noted include the stalk’s shape, its surface
texture, its colour, and the texture and colour of the context (the
flesh or internal tissue). If a ring (annulus) and basal cup (volva) are
present, they are described.
|
|
Microscopic Features
Spores
Spore
prints are key aids in identifying mushrooms. Separate a mature cap
from the stalk and place it on white or black paper with the hymenium
facing down, so that spores collect on the paper. Place the cap and
paper inside a lightly covered container and leave outside in ambient
temperatures for 12–24 hours. Note the spore print colour (without
magnification) to help identify and classify the mushroom. To examine
the spores microscopically, place the hymenium tissue on a microscope
slide in water or Melzer’s reagent and cover with a glass cover slip
before examining under
the microscope. Measure and examine the spores. Spores are placed in
Melzer’s reagent because the spores’ reaction helps in mushroom
identification. Sometimes, to identify unknown specimens, additional
reagents such as cotton blue are used as an aid to precise
identification.
Other Features
Clamp
connections, which are connecting cells between two adjoining hyphal
cells, occur only in fungi belonging to the Basidiomycetes. The presence
or absence of clamp connections can assist in identifying mushrooms in
this group.
The length of the basidia or asci and the number of spores produced by them also help to identify some mushrooms.
Hymenial
cystidia are sterile cells interspersed among the basidia in the
hymenium, and their presence or absence can also be important. When they
are present, their shape and size are very helpful in indentifying
mushrooms.
Mushroom Habitats and Habits
Habitat
includes the ecological setting in which a mushroom is most likely to
be found. This includes the host trees or host substrates as well as the
surrounding vegetation. Some mushrooms can colonize a wide range of
substrates such as dead wood or twigs or hosts, while others are very
specific to certain substrates or hosts. For instance, Boletus mirabilis, the admirable bolete, forms ectomycorrhizae only with western hemlock, while Tricholoma magnivelare,
the pine mushroom, colonizes a variety of conifer trees. Likewise, some
saprophytic mushrooms decompose a wide range of woody hosts, while
others decompose very specific woody hosts. Some saprophytes decompose
forest litter, while others decompose organic matter in pastures, lawns
or disturbed areas.
The
vast majority of B.C. NTFP mushrooms fruit in the autumn, although some
fruit only in the spring. Others may fruit throughout the year, weather
permitting. It is important to know in which season a mushroom fruits
when hunting for a specific mushroom. Fruiting itself is triggered by
such environmental factors as moisture and temperature. Fruiting
patterns of specific mushrooms vary, as some fruit singly, others fruit
in troops and still others form clusters. On this website, we provide
the fruiting habit of each mushroom.
Distribution Maps
The
distribution map for each species is based on collection areas
confirmed from personal and herbarium collections and published reports.
There are undoubtedly parts of the province where some species occur
but which are not yet included in these maps. As more data are added,
these distribution maps will more accurately reflect the entire range
and distribution of each species.
Interesting Facts
Mushrooms
have been part of human civilizations for centuries. One of the
earliest records of mushroom use was the case of Otzi, the Tyrolean
Iceman, who was frozen in ice but not discovered until 1991 in the
Italian Alps. Otzi lived 5,000 years ago and was carrying three
different mushrooms when he died. The fungi had laxative and antibiotic
properties, and scientists speculate that Otzi used the fungi to cure
his stomach parasites. In Asia, mushrooms have been used for centuries
for culinary, medicinal, nutriceutical and aphrodisiacal purposes.
Ancient Greeks and Romans welcomed thunderstorms because they believed
that mushrooms appeared after lightning. There is a plethora of
beneficial and unusual uses attributed to certain mushrooms, some of
which we include in our treatment of individual species. We also provide
unusual items of interest about each species. Some fungi have a
livelier or longer history than others, so for some species there is an
extensive section on interesting facts.
Disclaimer
No
single test can determine the culinary, beneficial or toxic properties
of a specific mushroom. As some mushrooms can be deadly poisonous, we
emphasize that the only safe means of separating edible mushrooms from
poisonous ones is the positive identification by an expert.
For
over twenty centuries, Chinese doctors have used mushrooms and other
fungi to cure many types of human diseases. These medicinal mushrooms
have also been credited with the ability to promote health and boost the
immune system. However, Western medicine is still struggling to confirm
Eastern medicinal uses of mushrooms.
Many
mushrooms produce both beneficial and toxic compounds; the same
compound can have beneficial or detrimental effects depending on the
amount consumed, how the mushroom is prepared, what other foods or
beverages are consumed with it, and whether an individual is
over-senstive to the active compounds. We neither confirm nor deny the
actions attributed to individual mushroom compounds. Although we are
aware of many incidences where people attribute their well being to a
specific mushroom, we try to avoid repeating anecdotal evidence and cite
only published scientific research with respect to the healing or other
beneficial properties attributed to each species. We do direct the
reader to published reports of the benefits of specific mushrooms after
each species description.
Reference
Wills,
R.M., and R.G. Lipsey, 1999. An economic strategy to develop Non-Timber
Forest Products and Services in British Columbia. Forest Renewal BC
Project No. PA97538-ORE. Final Report. |
|